Acari
	Eryiphyoidea (gall mites)
	Undetermined species causes galls on leaves.
Hemiptera
	Cicadellidae (leafhoppers)
	Require more research.
	Clastopteriadae (spittlebugs)
	Clastoptera sp. – A small sized spittlebug. Occasionally found on M. calvescens at the Turrialba sites. Host specificity remains unknown and will remain so until the taxonomy is worked out. V. Thompson (Roosevelt University, Chicago) and C. Godoy (INBio, Costa Rica) are contemplating working on the taxonomy of this genus, since as xylem feeders, they are potentially important vectors of an increasingly important bacterial disease (Xylosella, which affects a wide variety of plants and may be a reason for not using this insect in biological control, even if it proves host-specific).
	Membracidae (treehoppers)
	Micrutalis sp. (identified by C. Godoy) – Common on M. calvescens at various sites and times of year. Worth investigating.
	Psyllidae: Paurocephalinae
	Haplaphalara - 20 spp. in Costa Rica, principally on Melastomataceae. The genus name is being changed.
Coleoptera
	Curculionidae (weevils)
	Cryptorhynchus melastomae (identified by R. Anderson) – Apparently lays its eggs in cut stems. The larvae probably feed on damaged stems.
	Derelomini (identified by Jens).
	Exophthalmus jekelianus cf. (identified by Jens) – Adults feed on more or less mature leaves. Larvae are probably root-feeders, on undetermined plants.
	Pedetinus halticoides (Errirhinidae) (identified by Jens) – On fruits/seeds.
	Penestes (identified by Jens) – Said to do considerable damage to Clidemia hirta seedlings but larvae require succulent petiole.
Lepidoptera
	Arctiidae
	A species of Ctenuchinae, on leaves.
	Geometridae
	Undetermined species (apparently not in the INBio collection) – Larva feeds on mature leaves, Parasitoids: Euplectrus sp. (Eulophidae), 17 from a host larva, with dried host and cocoons of the Euplectrus; specimens sent to Christer Hansson.
	Riodinidae: Euselasiinae
	Euselasia – SW USA through South America, with the greatest number in the Amazon. Most species are rare and restricted to particular habitats. Many are crepuscular. Hostplants include Myrtaceae, Clusiaceae, Sapotaceae, and Melastomataceae. Eggs are of a fustrum shape, caterpillars are typically covered in downy setae, they may be solitay or gregarious and none have an association with ants. The gregarious larvae of some species move about in synchronous head-to-tail procession, feed at night, and rest during the day on the base of tree trunks. Pupate on top of the leaf. Adults rest with the wings folded over the back as in Lycaenidae. They visit a variety of small flowered plants and occasionally wet sand. Their activity is usually restricted to short windows of time during the day. Males of several species may use the same tree to perch on, but the species show a pronounced vertical stratification. Activity times of males and females may be very different. 27 spp. known from CR, hostplants known for 7: 2 from Miconia, 2 from Eugenia and 1 from Psidium (Myrtaceae), 1 from Clusia and 1 from Mammea (Clusiaceae). According to Burkhart, 2 species are very promising for controlling M.– Euselasia aurantiaca and E. chrysippe. They are voracious leaf feeders, but problematic in getting adults to feed or mate (probably require a large open sunlit area with vegetation around it to induce mating). We have found the latter but not the former; however we have found an additional species.
	E. bettina (identified by K. Nishida) - Larvae feed on mature leaves.
	E. chrysippe (identified by K. Nishida) - Larvae feed on mature leaves. Hosts: Miconia calvescens, M. elata. 0-1500m. Egg pale green, smooth. Larva pale yellow with 10 pairs of conspicuous black dorsal spots on the abdominal segments (and a pair on the third thoracic segment) that are divided by a dark midline; thoracic segments darker yellow than other body segments; all segments bear a sparse covering of long yellowish and black setae, with the black ones arising mainly from the dark dorsal spots and the yellowish setae arising laterally; head shiny amber with a sparse covering of setae. Processionary and synchronous in their feeding, movements, resting, and molting.
	Thecla opisena (identified by K. Nishida) - Larvae feed inside of fruits.
	Noctuidae
	Antiblemma sp. (VOB-904 INBio.) (identified by K. Nishida) - Larvae feed on small to mature leaves.
	Two other, unidentified species, feed on leaves (apparently not in the INBio collection.)
	Notodontidae
	Meragisa sp. 5 (=sp. 8 Schaus) (identified by K. Nishida) - Larvae feed on mature leaves.
	Oecophoridae
	Sp. 1 - on leaves.
	Sp. 2 - on leaves.
	Psychidae
	Undetermined species.
	Pterophoridae
	Undetermined species.
	Saturniidae
	Hylesia continua (identified by K. Nishida) - On leaves.

Download the summary presentation Miconia research in Costa Rica, given at The 2009 International Miconia Conference "Pulling it all Together".
May 4-7, 2009 Maui, Hawaii.

Authorship details
Hanson, P., 2003 (June 2). Natural Enemies in Insects.
In Biological Control of Miconia calvescens. http://www.miconia.net